A man buried in Russia in the Hunter-Gatherer era

The Hunter-Gatherer culture of the Samara region in Russia is a fascinating and foundational part of prehistoric human history. This era, which largely corresponds to the Mesolithic period, is characterized by the adaptation and survival strategies of Eurasian hunter-gatherers in the area around the Volga River basin. The period is significant due to its role in the transition from mobile foraging societies to more settled agricultural communities.
Geography and Environment
The Samara region is situated in the forest-steppe zone of the Middle Volga. This area provided a diverse and bountiful environment for hunter-gatherers. The landscape featured open steppes, dense forests, and numerous rivers and lakes, which offered a rich array of resources. The climate during this period was warming after the last glacial period, leading to the flourishing of flora and fauna that were crucial for sustenance.
Lifestyle and Subsistence
Hunter-gatherer communities of the Samara region were highly adaptive and mobile, exploiting the seasonal abundance of resources. Their subsistence strategies were diverse, including:
Hunting: The people primarily hunted large herbivores such as elk, deer, and boar. Hunting tools were crafted from bone and flint, demonstrating a high degree of skill in tool-making.
Fishing: Proximity to the Volga River and numerous smaller waterways made fishing a substantial component of their diet. They employed nets, hooks, and spears, taking advantage of the rich fish populations.
Gathering: The gathering of wild plants, berries, nuts, and tubers supplemented their diet. Seasonal availability dictated their movement and settlement patterns.
Social Structure and Organization
The social structure of Samara's hunter-gatherers was likely characterized by small, kin-based groups, with flexible membership and egalitarian principles. Leadership roles were probably situational and based on skill or knowledge rather than hereditary status.
The seasonal nature of their subsistence practices meant that these groups were semi-nomadic. They established temporary camps, which served as bases from which they could exploit nearby resources.
Material Culture and Technology
Their material culture was marked by the use of locally available materials such as flint, bone, and antler. The tools and artifacts created by these communities include:
Lithic Tools: These include blades, scrapers, and points, produced through sophisticated techniques such as pressure flaking.
Bone and Antler Tools: Implements such as needles, awls, and harpoons were carved, showcasing precision in craftsmanship.
Pottery: The Samara culture is significant for its early experimentation with pottery. While not as developed as later cultures, their pottery is some of the earliest in the region and represents a shift towards more sedentary lifestyles.
Rituals and Beliefs
There is evidence to suggest that the Samara hunter-gatherers engaged in ritual activities and had a complex belief system:
Burial Practices: Some burial sites indicate the deliberate placement of bodies and grave goods such as stone tools and ornaments, hinting at beliefs in an afterlife.
Art and Ornamentation: Personal adornments made from shells, animal teeth, and bones suggest not only a sense of aesthetic but also possible status indicators or ritualistic significance.
Genetic and Cultural Legacy
Genetically, the people of the Samara region were part of the broader genetic pool of Eurasian hunter-gatherers. They contributed to the genetic diversity of subsequent populations in Europe and Asia, especially during the Neolithic migrations and the later spread of Indo-European languages.
Culturally, the transition towards more complex technologies and social structures set the stage for the Neolithic Revolution, where agriculture and animal husbandry became prevalent. Insights into their adaptability, innovation, and social organization provide valuable understanding into the evolutionary trajectory of human societies in the region.
In conclusion, the hunter-gatherers of the Samara region in Russia played a critical role in the early chapters of human prehistory. Their ability to adapt to changing environments, innovate in tool-making, and develop rich cultural practices laid the groundwork for the complex societies that would follow.
Sample ID | Culture/Period | Date | Location | Action |
---|---|---|---|---|
I0234 | Srubnaya Culture | 1850 BCE | Samara Oblast. Volga River Valley. Samara Steppes. Rozhdestveno I, Russia | View |
I0126 | Middle Bronze Poltavka | 2867 BCE | Samara Oblast. Volga River Valley. Kutuluk River. Kutuluk III, Russia | View |
I0432 | Middle Bronze Poltavka | 2925 BCE | Samara Oblast. Sok River. Potapovka I, Russia | View |
I0434 | Eneolithic Khvalynsk Culture, Russia | 5198 BCE | Samara Oblast. Volga River Valley. Khvalynsk II, Russia | View |
I0433 | Eneolithic Khvalynsk Culture, Russia | 4697 BCE | Samara Oblast. Volga River Valley. Khvalynsk II, Russia | View |
I0122 | Eneolithic Khvalynsk Culture, Russia | 4936 BCE | Samara Oblast. Volga River Valley. Khvalynsk II, Russia | View |
I0124 | Hunter-Gatherer Samara, Russia | 5660 BCE | Samara Oblast. Volga River Valley. Sok River. Lebyazhinka, Russia | View |
I0370 | Early Bronze Samara Yamnaya | 3300 BCE | Samara Oblast. Volga River Valley. Eastern Orenburg. Pre-Ural steppe. Ishkinovka I, Russia | View |
I0441 | Early Bronze Samara Yamnaya | 3010 BCE | Samara Oblast. Volga River Valley. Buzuluk. Kurmanaevka III, Russia | View |
I0444 | Early Bronze Samara Yamnaya | 3335 BCE | Samara Oblast. Volga River Valley. Kutuluk River. Kutuluk, Russia | View |
I0439 | Early Bronze Samara Yamnaya | 3322 BCE | Samara Oblast. Volga River Valley. Sok River. Lopatino I, Russia | View |
I0357 | Early Bronze Samara Yamnaya | 3093 BCE | Samara Oblast. Volga River Valley. Sok River. Lopatino I, Russia | View |
I0429 | Early Bronze Samara Yamnaya | 3339 BCE | Samara Oblast. Volga River Valley. Sok River. Lopatino I, Russia | View |
I1282 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1302 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1276 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1284 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1274 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1280 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1314 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1277 | Chalcolithic Spain | 2570 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1272 | Chalcolithic Spain | 2857 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1281 | Chalcolithic Spain | 2867 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1300 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I0807 | Middle Neolithic Baalberge Culture, Germany | 3977 BCE | Esperstedt, Germany | View |
I0559 | Middle Neolithic Baalberge Culture, Germany | 3646 BCE | Quedlinburg Site IX, Germany | View |
I0560 | Middle Neolithic Baalberge Culture, Germany | 3637 BCE | Quedlinburg Site IX, Germany | View |
I1546 | Bell Beaker Culture, Germany | 2500 BCE | Benzingerode-Heimburg, Germany | View |
I0806 | Bell Beaker Culture, Germany | 2433 BCE | Quedlinburg Site VII, Germany | View |
I0805 | Bell Beaker Culture, Germany | 2465 BCE | Quedlinburg Site VII, Germany | View |
I0113 | Bell Beaker Culture, Germany | 2396 BCE | Quedlinburg Site XII, Germany | View |
I0112 | Bell Beaker Culture, Germany | 2455 BCE | Quedlinburg Site XII, Germany | View |
I1530 | Bell Beaker Culture, Germany | 2456 BCE | Rothenschirmbach, Germany | View |
I0111 | Bell Beaker Culture, Germany | 2474 BCE | Rothenschirmbach, Germany | View |
I0108 | Bell Beaker Culture, Germany | 2575 BCE | Rothenschirmbach, Germany | View |
I0171 | Late Neolithic Benzigerode-Heimburg, Germany | 2288 BCE | Benzingerode-Heimburg, Germany | View |
I0059 | Late Neolithic Benzigerode-Heimburg, Germany | 2343 BCE | Benzingerode-Heimburg, Germany | View |
I1542 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I1536 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I1544 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I1538 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I1539 | Corded Ware Culture, Germany | 2630 BCE | Esperstedt, Germany | View |
I0106 | Corded Ware Culture, Germany | 2461 BCE | Esperstedt, Germany | View |
I1540 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I1541 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I1532 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I0049 | Corded Ware Culture, Germany | 2463 BCE | Esperstedt, Germany | View |
I0211 | Hunter-Gatherer Karelia, Russia | 7050 BCE | Karelia. Yuzhnyy Oleni Ostrov, Russia | View |
I0061 | Hunter-Gatherer Karelia, Russia | 7050 BCE | Karelia. Yuzhnyy Oleni Ostrov, Russia | View |
I0550 | Late Neolithic Karsdorf, Germany | 2572 BCE | Karsdorf, Germany | View |
I0797 | Early Neolithic Linear Pottery Culture, Germany | 5500 BCE | Karsdorf, Germany | View |
I0795 | Early Neolithic Linear Pottery Culture, Germany | 5215 BCE | Karsdorf, Germany | View |
I0176 | Middle Neolithic Linear Pottery Culture, Hungary | 5209 BCE | Szemely-Hegyes, Hungary | View |
I0551 | Middle Neolithic Salzmuende Culture, Germany | 3400 BCE | Salzmünde-Schiepzig, Germany | View |
I0409 | Early Neolithic Spain | 5312 BCE | Els Trocs, Spain | View |
I0411 | Early Neolithic Spain | 5298 BCE | Els Trocs, Spain | View |
I0405 | Middle to Late Neolithic Spain | 3900 BCE | La Mina, Spain | View |
I0174 | Early Neolithic Starčevo Culture 1, Hungary | 5712 BCE | Alsonyek-Bataszek. Mérnöki telep, Hungary | View |
I0115 | Early Bronze Age Unetice Culture, Germany | 1959 BCE | Esperstedt, Germany | View |
I0117 | Early Bronze Age Unetice Culture, Germany | 2276 BCE | Esperstedt, Germany | View |
I0804 | Early Bronze Age Unetice Culture, Germany | 2139 BCE | Eulau, Germany | View |
I0803 | Early Bronze Age Unetice Culture, Germany | 2135 BCE | Eulau, Germany | View |
I0164 | Early Bronze Age Unetice Culture, Germany | 2027 BCE | Quedlinburg Site VIII, Germany | View |
I0114 | Early Bronze Age Unetice Culture, Germany | 2141 BCE | Esperstedt, Germany | View |
I0022 | Early Neolithic Linear Pottery Culture, Germany | 5500 BCE | Baden-Württemberg. Stuttgart-Mühlhausen. Viesenhaeuser Hof, Germany | View |
I0026 | Early Neolithic Linear Pottery Culture, Germany | 5500 BCE | Baden-Württemberg. Stuttgart-Mühlhausen. Viesenhaeuser Hof, Germany | View |
I0013 | Hunter-Gatherer Motala, Sweden | 5966 BCE | Motala. Kanaljorden, Sweden | View |
I0011 | Hunter-Gatherer Motala, Sweden | 5721 BCE | Motala. Kanaljorden, Sweden | View |
I0015 | Hunter-Gatherer Motala, Sweden | 5967 BCE | Motala. Kanaljorden, Sweden | View |
I0012 | Hunter-Gatherer Motala, Sweden | 5715 BCE | Motala. Kanaljorden, Sweden | View |
I0014 | Hunter-Gatherer Motala, Sweden | 5885 BCE | Motala. Kanaljorden, Sweden | View |
I0017 | Hunter-Gatherer Motala, Sweden | 5722 BCE | Motala. Kanaljorden, Sweden | View |
I1508 | Early Neolithic Körös Culture, Hungary | 5716 BCE | Berettyóújfalu-Morotva-Liget, Hungary | View |
I1500 | Middle Neolithic Alföld Linear Pottery Culture, Hungary | 5301 BCE | Kompolt-Kigyoser, Hungary | View |
I1100 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1102 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1099 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1103 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1101 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1097 | Neolithic Turkey | 6420 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0744 | Neolithic Turkey | 6400 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1096 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1098 | Neolithic Turkey | 6419 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0708 | Neolithic Turkey | 6224 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0745 | Neolithic Turkey | 6387 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0746 | Neolithic Turkey | 6070 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0707 | Neolithic Turkey | 6225 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0709 | Neolithic Turkey | 6223 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0725 | Neolithic Turkey | 6400 BCE | Menteşe, Turkey | View |
I0736 | Neolithic Turkey | 6500 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1499 | Middle Neolithic Alföld Linear Pottery Culture Bükk Group, Hungary | 5286 BCE | Garadna, Hungary | View |
I1502 | Early Bronze Age Makó Culture, Hungary | 2195 BCE | Kompolt-Kigyoser, Hungary | View |
I1497 | Late Chalcolithic Baden Culture, Hungary | 3320 BCE | Apc-Berekalya I, Hungary | View |
I1495 | Late Neolithic Lengyel Culture, Hungary | 4496 BCE | Apc-Berekalya I, Hungary | View |
I1498 | Middle Neolithic Alföld Linear Pottery Culture, Hungary | 5297 BCE | Debrecen Tocopart Erdoalja, Hungary | View |
I1507 | Early Neolithic Hunter-Gatherer Körös Culture, Hungary | 5788 BCE | Tiszaszolos-Domaháza, Hungary | View |
I1496 | Middle Neolithic Linear Pottery Culture, Hungary | 5211 BCE | Apc-Berekalya I, Hungary | View |
I1505 | Middle Neolithic Alföld Linear Pottery Culture, Hungary | 5210 BCE | Polgár-Ferenci hát. M3-31, Hungary | View |
I1271 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1303 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I1549 | Bell Beaker Culture, Germany | 2500 BCE | Benzingerode-Heimburg, Germany | View |
I0407 | Middle to Late Neolithic Spain | 3900 BCE | La Mina, Spain | View |
I0025 | Early Neolithic Linear Pottery Culture, Germany | 5500 BCE | Baden-Württemberg. Stuttgart-Mühlhausen. Viesenhaeuser Hof, Germany | View |
I0247 | Questionable Iron Age Scythian, Russia | 385 BCE | Samara Oblast. Volga River Valley. Volga Steppes. Nadezhdinka, Russia | View |
I0423 | Srubnaya Culture | 1850 BCE | Samara Oblast. Volga River Valley. Samara River. Barinovka I, Russia | View |
I0443 | Early Bronze Samara Yamnaya | 3300 BCE | Samara Oblast. Volga River Valley. Sok River. Lopatino II, Russia | View |
I0726 | Neolithic Turkey | 6400 BCE | Menteşe, Turkey | View |
I0103 | Corded Ware Culture, Germany | 2617 BCE | Esperstedt, Germany | View |
I0172 | Middle Neolithic Esperstedt, Germany | 3363 BCE | Esperstedt, Germany | View |
I0408 | Middle to Late Neolithic Spain | 3895 BCE | La Mina, Spain | View |
I0412 | Early Neolithic Spain | 5309 BCE | Els Trocs, Spain | View |
I0054 | Early Neolithic Linear Pottery Culture, Germany | 5216 BCE | Unterwiederstedt, Germany | View |
I0104 | Corded Ware Culture, Germany | 2563 BCE | Esperstedt, Germany | View |
I0116 | Early Bronze Age Unetice Culture, Germany | 2137 BCE | Esperstedt, Germany | View |
I0118 | Late Neolithic Alberstedt, Germany | 2469 BCE | Alberstedt, Germany | View |
I0406 | Middle to Late Neolithic Spain | 3900 BCE | La Mina, Spain | View |
I0410 | Early Neolithic Spain | 5298 BCE | Els Trocs, Spain | View |
I0413 | Early Neolithic Spain | 5304 BCE | Els Trocs, Spain | View |
I0235 | Srubnaya Culture | 1850 BCE | Samara Oblast. Volga River Valley. Samara Steppes. Rozhdestveno I, Russia | View |
I0374 | Middle Bronze Poltavka | 2800 BCE | Samara Oblast. Volga River Valley. Samara River. Nikolaevka III, Russia | View |
I1583 | Neolithic Turkey | 6424 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0581 | Chalcolithic Spain | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View |
I0440 | Middle Bronze Poltavka | 2887 BCE | Samara Oblast. Volga River Valley. Sok River. Lopatino II, Russia | View |
I0371 | Middle Bronze Poltavka | 2871 BCE | Samara Oblast. Volga River Valley. Sok River. Grachevka, Russia | View |
I0438 | Early Bronze Samara Yamnaya | 3020 BCE | Samara Oblast. Volga River Valley. Samara River. Luzkhi I, Russia | View |
I0418 | Middle Bronze Poltavka | 2131 BCE | Samara Oblast. Volga River Valley. Samara River. Utyevka VI, Russia | View |
I0099 | Late Bronze Age Halberstadt, Germany | 1202 BCE | Halberstadt-Sonntagsfeld, Germany | View |
I0047 | Early Bronze Age Unetice Culture, Germany | 2116 BCE | Halberstadt-Sonntagsfeld, Germany | View |
I1534 | Corded Ware Culture, Germany | 2500 BCE | Esperstedt, Germany | View |
I0060 | Bell Beaker Culture, Germany | 2404 BCE | Rothenschirmbach, Germany | View |
I0723 | Neolithic Turkey | 6008 BCE | Menteşe, Turkey | View |
I0724 | Neolithic Turkey | 6400 BCE | Menteşe, Turkey | View |
I0727 | Neolithic Turkey | 6400 BCE | Menteşe, Turkey | View |
I1580 | Neolithic Turkey | 6381 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1581 | Neolithic Turkey | 6386 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1585 | Neolithic Turkey | 6217 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I1579 | Neolithic Turkey | 6221 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0854 | Neolithic Turkey | 6228 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View |
I0581 | 2900 BCE | Burgos. Atapuerca. El Mirador Cave, Spain | View | |
I0440 | 2887 BCE | Samara Oblast. Volga River Valley. Sok River. Lopatino II, Russia | View | |
I0418 | 2131 BCE | Samara Oblast. Volga River Valley. Samara River. Utyevka VI, Russia | View | |
I0438 | 3020 BCE | Samara Oblast. Volga River Valley. Samara River. Luzkhi I, Russia | View | |
I0585 | 5982 BCE | Leon. La Brana-Arintero, Spain | View | |
I1534 | 2500 BCE | Esperstedt, Germany | View | |
I1504 | 987 BCE | Ludas-Varjú dűlő, Hungary | View | |
I0099 | 1202 BCE | Halberstadt-Sonntagsfeld, Germany | View | |
I0047 | 2116 BCE | Halberstadt-Sonntagsfeld, Germany | View | |
I1579 | 6221 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View | |
I1581 | 6386 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View | |
I1580 | 6381 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View | |
I1585 | 6217 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View | |
I1583 | 6424 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View | |
I0727 | 6400 BCE | Menteşe, Turkey | View | |
I0724 | 6400 BCE | Menteşe, Turkey | View | |
I0854 | 6228 BCE | Northwest Anatolia. Marmara. Barcın, Turkey | View | |
I0723 | 6008 BCE | Menteşe, Turkey | View | |
I0371 | 2871 BCE | Samara Oblast. Volga River Valley. Sok River. Grachevka, Russia | View | |
I0060 | 2404 BCE | Rothenschirmbach, Germany | View |
Sample ID | Culture/Period | Date | Location | Action |
---|---|---|---|---|
I0124 | Hunter-Gatherer Samara, Russia | 5660 BCE | Samara Oblast. Volga River Valley. Sok River. Lebyazhinka, Russia | View |
Ancient genetic admixture analysis compares the DNA profile of this individual (I0124) with present-day reference populations. These results show what percentage of the individual's genetic makeup resembles ancient populations from different geographic regions.
Modern genetic admixture analysis compares the DNA profile of this individual (I0124) with present-day reference populations. These results show what percentage of the individual's genetic makeup resembles modern populations from different geographic regions.
These results complement the ancient ancestry components shown in the previous section, offering a different perspective on the individual's genetic profile by comparing it with modern reference populations rather than prehistoric ancestral groups.
The G25 coordinates for the sample I0124 are as follows. You can analyze its admixture using G25 Studio.
I0124,0.11986606,0.03409,0.08182972,0.0667483,0.0059761,0.01313468,-0.00419378,-0.0080473,-0.00905372,-0.03274804,0.00890576,-0.0099419,0.01565672,0.0058434,-0.01419766,-0.01943738,-0.01240572,-0.0014921,-0.00997694,-0.00794618,-0.0040387,0.004441,-0.00964554,0.00187796,-0.00449308
Genome-wide patterns of selection in 230 ancient Eurasians
Ancient DNA makes it possible to observe natural selection directly by analysing samples from populations before, during and after adaptation events. Here we report a genome-wide scan for selection using ancient DNA, capitalizing on the largest ancient DNA data set yet assembled: 230 West Eurasians who lived between 6500 and 300 bc, including 163 with newly reported data. The new samples include, to our knowledge, the first genome-wide ancient DNA from Anatolian Neolithic farmers, whose genetic material we obtained by extracting from petrous bones, and who we show were members of the population that was the source of Europe's first farmers. We also report a transect of the steppe region in Samara between 5600 and 300 bc, which allows us to identify admixture into the steppe from at least two external sources. We detect selection at loci associated with diet, pigmentation and immunity, and two independent episodes of selection on height.